Ray Romano

as Manny (voice)

John Leguizamo

as Sid (voice)

Denis Leary

as Diego (voice)

Seann William Scott

as Crash (voice)

Josh Peck

as Eddie (voice)

Queen Latifah

as Ellie (voice)

Jennifer Lopez

as Shira (voice)

Peter Dinklage

as Captain Gutt (voice)

Wanda Sykes

as Granny (voice)

Keke Palmer

as Peaches (voice)

Josh Gad

as Louis (voice)

Nick Frost

as Flynn (voice)

Aziz Ansari

as Squint (voice)

Alain Chabat

as Silas (voice)

Kunal Nayyar

as Gupta (voice)

Drake

as Ethan (voice)

Nicki Minaj

as Steffie (voice)

Rebel Wilson

as Raz (voice)

Alan Tudyk

as Milton / Hunky Siren (voice)

Ester Dean

as Female Siren (voice)

Eddie 'Piolin' Sotelo

as Uncle Fungus (voice)

Joy Behar

as Eunice (voice)

Heather Morris

as Katie (voice)

Chris Wedge

as Scrat (voice)

Christopher Campbell

as Creature Siren (voice)

Karen Disher

as Scratte (voice)

Marieve Herington

as Additional Voices (voice)

Lucas Leguizamo

as Additional Voices (voice)

Allegra Leguizamo

as Additional Voices (voice)

Jason Fricchione

as Dumb Mammoth / Various (voice)

Ben Gleib

as Marshall (voice)

George Jacobs

as Beaver (voice)

James Palumbo

as Panicked Start (voice)

Simon Pegg

as Buck (voice)

Alexandra Romano

as Meghan (voice)

Patrick Stewart

as Ariscratle (voice)

Brian Scott McFadden

as Additional Voices (voice)

Ashley Albert

as Additional Voices (voice)

Edita Brychta

as Additional Voices (voice)

Dennis T. Carnegie

as Additional Voices (voice)

Aimée Castle

as Additional Voices (voice)

Jim Conroy

as Additional Voices (voice)

Jason Harris

as Additional Voices (voice)

Sondra James

as Additional Voices (voice)

Alexa Kahn

as Additional Voices (voice)

Bill Lobley

as Additional Voices (voice)

Chris Phillips

as Additional Voices (voice)

Tara Sands

as Additional Voices (voice)

Keith Silverstein

as Additional Voices (voice)

Vanessa Lemonides

as Additional Voices (voice)

Danny Flaherty

as Additional Voices (voice)

Linda Joy

as Additional Voices (voice)

Selenis Leyva

as Additional Voices (voice)

Ron McClary

as Additional Voices (voice)

Peter Pamela Rose

as Additional Voices (voice)

Gabriel Rush

as Additional Voices (voice)

Randy Thom

as Additonal Voices (voice)

Annie Maginnis Tippe

as Additional Voices (voice)

Corwin C. Tuggles

as Additional Voices (voice)

Tashiana Washington

as Additional Voices (voice)

Frank Welker

as Giant Crab (voice) (uncredited)