Warwick Davis

as Master Moley (voice)

Gemma Arterton

as Mona Lisa (voice)

Togo Igawa

as Mystic Mole (voice)

Richard E. Grant

as The Gardener (voice)

Charles Dance

as Mr. James Moley (voice)

Julie Walters

as Mrs. Elizabeth Moley / Queen (voice)

Calum Callaghan

as Milo / American Elder (voice)

Amanda Drew

as Melony (voice)

Sean Baker

as Royal Guard / Guest (voice)

James Reatchlous

as The Duke (voice)

Tony Nottage

as Geeky Mole (voice)

Arthur Landon

as Rodrigo (voice)

Leia Nottage

as Child Mole (voice)

Kris Milnes

as (voice)

Joseph Balderrama

as (voice)

Stephen Critchlow

as (voice)

Rupert Holliday-Evans

as (voice)

Jo Hall

as (voice)

Stephanie Thomas

as (voice)

Alison Dowling

as (voice)

Julia Brans

as (voice)